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1、PBG650AdvancedPlantBreedingModule13:BreedingforDiverseEnvironmentsGenotypexEnvironmentInteractionsGenotypebyenvironmentinteractionGenotypesresponddifferentlyacrossarangeofenvironmentsi.e.,therelativeperformanceofvarietiesdependsontheenvironmentGXE,GEI,GbyE,GEGe
2、notypebyenvironmentinteractionsarecommonformostquantitativetraitsofeconomicimportanceAdvancedbreedingmaterialsmustbeevaluatedinmultiplelocationsformorethanoneyearMET=multi-environmenttrialsP=G+E+GETypesofGEIInteractionmaybedueto:heterogeneityofgenotypicvariance
3、acrossenvironmentsimperfectcorrelationofgenotypicperformanceacrossenvironmentsnoncrossovercrossoverTheGEIchallengeEnvironmentaleffectisthegreatest,butisirrelevanttoselection(remember70-20-10rule,E:GE:G)Manystatisticalapproachesconsiderallofthephenotypicvariatio
4、n(i.e.,meansacrossenvironments),whichmaybemisleadingNeedanalysesthatwillhelpyoutocharacterizeGEI"GEInteractionisnotmerelyaproblem,itisalsoanopportunity"(Simmonds,1991).SpecificadaptationscanmakethedifferencebetweenagoodvarietyandasuperbvarietyTheGEIchallengeSom
5、eenvironmentalvariationispredictablecanbeattributedtospecific,characteristicfeaturesoftheenvironmente.g.,soiltype,soilfertility,plantdensitySomevariationisunpredictablee.g.,rainfall,temperature,humidityQuestionsabreedermayaskHowdoIchoosesuitableenvironmentsfort
6、esting?HowdoIallocateresources(numberoftestingsitesvsreplicateswithinsites)?Ingeneral,increasingenvironmentsdoesmoretoreducethestandarderrorofgenotypemeansthanincreasingreplicationsConventionalANOVARatiosofSumsofSquares(SS)canbeusedtoestimatethecontributionofin
7、teractionstothetotalSSCanalsoestimatevariancecomponentsduetoGXEMixedModelsforanalysisofMETForevaluationofGEIamonganelitegroupofcultivars,genotypesareoftenconsideredtobefixedeffectsandenvironmentsarerandom.TheGEIcomponentisrandom.OnecanobtainBLUPestimatesofGEIef
8、fects.ForthepurposeofestimatingbreedingvaluesusingBLUP,genotypesareconsideredtoberandomandenvironmentsarefixedeffects.Somestatisticiansbelievethatgenotypesshouldalwaysberand